3 down
Host,
part
of
TV
duo,
is
a
crawler
(4,3)
Host, part of TV duo, is a crawler (4,3)
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The definition is at the end of the clue.
How the clue works
Right, so "a crawler" at the end is the definition — army ants do exactly that, crawling about in their columns. Before that, "host" gives you ARMY (as in a host or multitude of something), and "part of TV duo" points to ANT, half of the telly pairing Ant and Dec. Stick ARMY and ANT together and you're there.