6 down
South
American
vehicle
in
front
of
North
American
ecosystem
(7)
South American vehicle in front of North American ecosystem (7)
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The definition is at the end of the clue.
How the clue works
Look at ecosystem at the end — that's your definition, the grassy plain. You build the answer by stringing together SA (South American), VAN (vehicle), and NA (North American) one in front of the other.